If you have a system with multiple power supplies and all the components connected to eachother in some way, it is important to make sure that all the voltages start from the same place. Remember that the voltage is actually the potential difference and that voltage is measured with respect to something. This "something" should be the same thing across the system.
As an analogy, think about building a house. The door frame may be 6'3" high but if your floor is 6" thick, the doorway will be too small - or will it? It depends where you are measuring from. So you define a "ground level" before you start building and measure everything up (or down) from this reference point.
In the same way, you link the negatives of all the power supplies together to define the "ground level" for all voltage measurements.
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